*Hatsumi Nishikawa¹, Humio Mitsudera¹, Takeshi Okunishi², Shin-ichi Ito³, Taku Wagawa⁴, Daisuke Hasegawa², Toru Miyama⁵, Hitoshi Kaneko² (1.Institute of Low Temperature Science, Hokkaido University, 2.Tohoku National Fisheries Research Institute, Japan Fisheries Research and Education Agency, 3.Atmosphere and Ocean Research Institute, The University of Tokyo, 4.Japan Sea National Fisheries Research Institute, Japan Fisheries Research and Education Agency, 5.Japan Agency for Marine-Earth Science and Technology)