*Xiaoyi Zeng1, Michael Boyle1, Runxi Wang1, David Baker1, Jonathan Cybulski2, Billy Hau1, Timothy Bonebrake1, Hannah Griffiths3, Paul Eggleton4, Louise Ashton1
(1. University of Hong Kong (Hong Kong), 2. Smithsonian Tropical Research Institute (Panama), 3. University of Bristol (UK), 4. Natural History Museum (UK))