Liyang Ma1,2, Zhilang Li1,2, Guanlin Li1, Wentong Jia1,2, Yanlei Liu1,2, Yuxia Li1, Yanling Wang1,2(1. State Key Laboratory of Stem cell and Reproductive Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China, 2. University of the Chinese Academy of Sciences, Beijing, China)