*Julie - An N. Gregorio^1,2、Elfritzson M. Peralta^3,5、Irisse Bianca B. de Jesus^3,5、Tohru Ikeya⁶、Jonathan Carlo A. Briones^3,4,5、Rey Donne S. Papa^3,4,5、Francis S. Magbanua¹、Noboru Okuda⁶ (1.Institute of Biology, University of the Philippines, Diliman, Quezon City, 1101 Philippines、2.Institute of Environmental Science and Meteorology, University of the Philippines, Diliman, Quezon City 1101 Philippines、3.Research Center for the Natural and Applied Sciences, University of Santo Tomas, España Boulevard, Manila, 1015, Philippines、4.Department of Biological Sciences, University of Santo Tomas, España Boulevard, Manila, 1015, Philippines、5.The Graduate School, University of Santo Tomas, España Boulevard, Manila, 1015, Philippines、6.Research Institute for Humanity and Nature, 457-4 Motoyama, Kamigamo, Kyoto, 603-8047, Japan)