*Aleksandr Nogovitcyn¹、Atsuko Sugimoto²、Ruslan Shakhmatov¹、Tomoki Morozumi¹、Shunsuke Tei²、Yumiko Miyamoto²、Trofim Maximov^3,4 (1.Graduate School of Environmental Science, Hokkaido University, Sapporo, Hokkaido, Japan、2.Arctic Research Center, Hokkaido University, Sapporo, Hokkaido, Japan、3.Institute for Biological Problems of Cryolithozone, Siberian Blanch of Russian Academy of Sciences, Yakutsk, Yakutia, Russia、4.International Center for Biogeoscience Educational and Scientific Trainings (BEST), North-Eastern Federal University, Yakutsk, Yakutia, Russia)