*Segawa Takahiro¹, Ryo Matsuzaki², Ayumi Akiyoshi³, Takahiro Yonezawa⁴, Hiroshi Mori⁵, Nozomu Takeuchi⁶ (1.University of Yamanashi, 2.National Institute for Environmental Studies, 3.National Institute of Polar Research, 4.Tokyo University of Agriculture, 5.National Institute of Genetics, 6.Chiba University)