*Hiroyuki Muraoka¹, Yayoi Takeuchi², Takehisa Yamakita³, Yuichi Kano⁴, Shin Nagai³, Sheila Vergara⁵, Eun-Shik Kim⁶, Tetsukazu Yahara^4,7 (1.Gifu University, 2.National Institute for Environmental Studies, 3.Japan Agency for Marin-Earth Science and Technology, 4.Kyusyu University, 5.ASEAN Centre for Biodiversity, 6.Kookmin University, 7.Kyusyu Open University)