*Salla A. M. Tenhovirta^1,2、Lukas Kohl^1,2,3、Markku Koskinen^1,2、Tatu Polvinen^1,2、Mari Pihlatie^1,2,4 (1.Environmental Soil Science, Department of Agricultural Sciences, University of Helsinki, Finland、2.Institute for Atmospheric and Earth System Research / Forest Sciences, University of Helsinki, Finland、3.Department of Environmental and Biological Sciences, University of Eastern Finland, Kuopio, Finland、4.Viikki Plant Science Centre (ViPS), University of Helsinki, Finland)