*Dai Yanagihara^1,2, Meiko Asaka^2,1, Takumi Komatsu¹, Di Wang^3,4, Shinya Honda³, Kei Watase⁵, Hidehiro Mizusawa^5,6, Kinya Ishikawa⁴, Shigeomi Shimizu³ (1. Graduate School of Arts and Sciences, The University of Tokyo, Tokyo, Japan, 2. Cognition and Behavior Joint Research Laboratory, RIKEN Center for Brain Science, Saitama, Japan, 3. Medical Research Institute, Tokyo Medical and Dental University, Tokyo, Japan, 4. Graduate School of Medical and Dental Sciences, Tokyo Medical and Dental University, Tokyo, Japan, 5. Center for Brain Integration Research, Tokyo Medical and Dental University, Tokyo, Japan, 6. National Center of Neurology and Psychiatry, Tokyo, Japan)