*Misato Kikuchi¹, Takuya Okada^1,2, Ken Miya², Kazuko Keino-Masu^1,2, Kenta Kobayashi³, Masayuki Masu^1,2 (1. Doctoral Program in Medical Sciences, Graduate School of Comprehensive Human Sciences, University of Tsukuba, Tsukuba, Japan, 2. Department of Molecular Neurobiology, Division of Biomedical Science, Institute of Medicine, University of Tsukuba, Tsukuba, Japan, 3. Section of Viral Vector Development, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, National Institutes of Natural Sciences, Okazaki, Japan)