*Kei Oyama^1,4, Yuji Nagai¹, Yukiko Hori¹, Haruhiko Iwaoki¹, Yuki Hori¹, Toshiyuki Hirabayashi¹, Mark A G Eldridge², Junya Hirokawa¹, Taku Banno¹, Ken-ichi Inoue³, Richard C Saunders², Masahiko Takada³, Makoto Higuchi¹, Barry J Richmond², Takafumi Minamimoto¹ (1. Advanced Neuroimaging Center, QST, Chiba, Japan, 2. Laboratory of Neuropsychology, National Institute of Mental Health, National Institutes of Health, Bethesda, USA, 3. Systems Neuroscience Section, Primate Research Institute, Kyoto University, Inuyama, Japan, 4. PRESTO, Japan Science and Technology Agency, Saitama, Japan)