*Sihui Cheng¹, Qirun Wang², Guge Suri³, Chenglin Miao⁴, Jie Lin² (1. Peking-Tsinghua Center for Life Sciences, Peking University, Beijing, China, 2. Center for Quantitative Biology, Peking University, Beijing, China, 3. PKU-IDG/McGovern Institute for Brain Research, Peking University, Beijing, China, 4. School of Life Sciences, Peking University, Beijing, China)