*Emi Fukuzawa¹, Kazuki SHIN'YA^1,2, Yuki Ito¹, Tomohiro Miyasaka³, Nobuto Kakuda¹, Harada Akihiro⁴, Kohta I Kobayashi¹ (1. Doshisha University , 2. JSPS Research Fellowship for Young Scientists, 3. Nihon University, 4. Osaka University)