*Kosuke Okuya¹, Reiko Yoshida¹, Rashid Manzoor¹, Shinji Saito², Tadaki Suzuki³, Osamu Ichii⁴, Hideaki Higashi¹, Ayato Takada¹ (1. Research Center for Zoonosis Control, Hokkaido University, 2. Influenza Virus Research Center, National Institute of Infectious Diseases, 3. Department of Pathology, National Institute of Infectious Diseases , 4. Faculty of Veterinary Medicine, Hokkaido University)

*Tomo Daidoji¹, Junichi Kajikawa¹, Yasuha Arai¹, Yohei Watanabe¹, Takaaki Nakaya¹ (1. Kyoto Prefectural University of Medicine)

*Kouji SAKAI¹, Naoya Nagata¹, Kimie Nomura², Takashi Irie³, Kazuaki takehara⁴, Hiroaki Kanda², Yasushi Ami¹ (1. National Institute of Infectious Diseases, 2. Department of Pathology, Saitama Cancer Center, 3. Department of Virology, Graduate School of Biomedical & Health Sciences, Hiroshima University, 4. Department of Veterinary Medicine, Tokyo University of Agriculture and Technology)

Yuto Kikutani¹, *Masatoshi Okamatsu¹, Shoko Nishihara², Sayaka Takase-Yoden², Takahiro Hiono¹, Robert P de Vries³, Ryan McBride⁴, Keita Matsuno^1,5, Hiroshi Kida^5,6, Yoshihiro Sakoda^1,5 (1. Faculty of Veterinary Medicine, Hokkaido University, 2. Soka University, 3. Utrecht Univerisity, 4. The Scripps Research Institute, 5. GI-CoRE, Hokkaido University, 6. Research Center for Zoonosis Control, Hokkaido University)