*Tetsuro AGUSA¹, Takashi KUNITO², Suguru INOUE¹, Tu Binh Minh³, Nguyen Minh Tue¹, Nguyen Ngoc Ha⁴, Nguyen Phuc Cam Tu⁴, Pham Thi Kim Trang³, Shin TAKAHASHI⁵, Bui Cach Tuyen⁶, Pham Hung Viet³, Hisato IWATA¹, Shinsuke TANABE¹ (1.Center for Marine Environmental Studies (CMES), Ehime University, Japan, 2.Faculty of Science, Shinshu University, Japan, 3.Hanoi University of Science, Vietnam National University, Vietnam, 4.Faculty of Fisheries, Nong Lam University, Vietnam, 5.Faculty of Agriculture, Ehime University, Japan, 6.Research Institute for Biotechnology and Environment (RIBE), Nong Lam University, Vietnam)