*Kohki Sowa¹, Kentaro Tanaka², Tsuyoshi Watanabe³, Jens Zinke⁴, Keiichi Nomura⁵, Malcolm McCulloch ^6,7 (1.Japan Agency for Marine-Earth Science and Technology, 2.Atmosphere and Ocean Research Institute, The University of Tokyo, 3.Hokkaido University, 4.Freie Universität Berlin , 5.Kushimoto Marine Park Center Ltd., 6.The University of Western Australia, 7.ARC Centre of Excellence for Coral Reef Studies)