*Ueno Hiromichi¹, Ryo Dobashi², Yuki Okada², Takahiro Tanaka³, Jun Nishioka⁴, Toru Hirawake¹, Atsushi Ooki¹, Sachihiko Itoh⁵, Yoshikazu Sasai⁶, Hideharu Sasaki⁶, Ichiro Yasuda⁵ (1.Faculty of Fisheries Sciences, Hokkaido University, 2.Graduate School of Fisheries Sciences, Hokkaido University, 3.Tohoku National Fisheries Research Institute, Japan Fisheries Researh and Education Agency, 4.Institute of Low Temperature Science, Hokkaido University, 5.Atmosphere and Ocean Research Institute, The University of Tokyo, 6.Japan Agency for Marine-Earth Science and Technology)