*Kanae Kobayashi¹, Keitaro Fukushima², Yuji Onishi², Kazuya Nishina³, Akiko Makabe⁴, Mamoru Oshiki⁵, Keisuke Koba², Satoshi Okabe¹ (1.Hokkaido Univ., 2.Center for Ecological Research, Kyoto Univ., 3.National Institute for Environmental Studies, 4.Japan Agency for Marine-Earth Science and Technology, 5.NIT, Nagaoka Coll.)