*Kamille Nacalaban Poblete¹、Dimple May Gianne Q. Dumaguit¹、Elfritzson M. Peralta^1,2,3、Francis S. Magbanua⁴、Rey Donne S. Papa^1,2,3、Noboru Okuda^5,6 (1.Department of Biological Sciences, College of Science, University of Santo Tomas, España Boulevard, Manila, 1015, Philippines、2.The Graduate School, University of Santo Tomas, España Boulevard, Manila, 1015, Philippines、3.Research Center for the Natural and Applied Sciences, University of Santo Tomas, España Boulevard, Manila, 1015, Philippines、4.Institute of Biology, University of the Philippines Diliman, Quezon City, 1101, Philippines、5.Research Center for Inland Seas, Kobe University, Japan、6.Research for Humanity & Nature, Japan)