*Chris Leong¹、Daiki Tomojiri¹、Ryuichi Shinjo^1,3、Jun Yasumoto²、Bam Razafindrabe² (1.Research Institute for Humanity and Nature, Kyoto, Japan、2.Faculty of Agriculture, University of the Ryukyus, Okinawa, Japan、3.Faculty of Science, University of the Ryukyus, Okinawa, Japan)