*Hiroaki Ohishi¹, Soya Shinkai², Hitoshi Owada³, Takeru Fujii¹, Kazufumi Hosoda⁴, Shuichi Onami², Takashi Yamamoto³, Yasuyuki Ohkawa¹, Hiroshi Ochiai¹ (1. Kyushu Univ. MIB, 2. RIKEN BDR, 3. Hiroshima Univ. Graduate School of Integrated Sciences for Life, 4. Ansanga Lab)

*Kaidi Fan^2,1, Leyan Cao ^1,2, Gideon Kipkemoi^1,2, Wenhao Wang ^1,2, Haruko Takahashi^1,2, Yutaka Kikuchi^1,2 (1. Graduate School of Integrated Sciences for Life, Hiroshima University, 2. Department of Biological Science, Graduate School of Science, Hiroshima University)

Yuki Sato¹, *Makoto T Hayashi^2,3 (1. Graduate School of Biostudies, Kyoto University, 2. IFOM-KU JRL, Graduate School of Medicine, Kyoto University, 3. Chromosome Instabilities Unit, IFOM ETS)

*Yasuhiro Hirano^1,2, Tatsuo Fukagawa¹ (1. Graduate School of Frontier Biosciences, Osaka University, 2. The Cancer Institute)

*Akio Nishimoto¹, suetake isao², yamada mio³, yosida tomomi³, toshiki takei¹, sinndou hideo³, hozyo hironobu¹ (1. Osaka University, 2. Nakamura Gakuen University, 3. National Center for Global Health and Medicine)

*Yohei Kono¹, Yixian Zheng², Stephen A. Adam³, Robert D. Goldman³, Hiroshi Kimura⁴, Takeshi Shimi¹ (1. Nano Life Science Institute, Kanazawa University, 2. Carnegie Institution for Science, 3. Northwestern University, 4. Cell Biology Center, Tokyo Institute of Technology)